After their adventures on The Earth That Was, the crew of the Serenity are reunited under the leadership of Captain Kaylee. Unfortunately with food, credits, and morale low, the crew takes on a heist which proves too much for their moral code and quickly turns into a recovery mission. And it's not long before their current dilemma puts Captain Kaylee, Mal, Zoe, Simon, River, Leonard, and Emma on a collision course with the mysterious past of the crew member who dragged them into this mess in the first place. That's right, the Hero of Canton himself, one Jayne Cobb.